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As seen in Figure 8B, unlike ADAM12, expression of ADAM19 and 33 (individually) also inhibited α5β1-mediated cell migration. As seen in Figure 8A, expression of both ADAM19 and ADAM33 inhibited α4β1-mediated cell migration. A higher expression level (4 μg per 6-cm dish) of ADAM33 inhibited α5β1-mediated cell migration, but its effect on α4β1-mediated migration was diminished (Huang, Bridges, and White, unpublished data). (B) Scratch wound assays were repeated as in panel A, except that α5-mediated cell migration was scored on the CCBD region of FN. Thus, strikingly, ADAM12 and ADAM17 demonstrate reciprocal effects on α4β1 and α5β1 mediated cell migration (Figure 8, A and B). Collectively, the results of Figures 5 and 6 show that the inhibitory effect of ADAM12 on α4β1-mediated cell migration does not require that it be proteolytically processed nor that it be proteolytically active.
Possible Mechanisms by Which ADAMs Influence Integrin-mediated Cell Migration
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